西藏豆科植物区系的形成与分化

 Xizang (Tibet) is rich in Leguminosae flora, comprising 41 genera and 254 species so far known, exclusive of the commonly cultivated taxa (including 11 genera and 16 species). There are 4 endemic genera (with 8 species), 10 temperate genera (with 175 species) and 19 tropical genera (with 46 species) as well as the representatives of those genera whose distribution centers are in East Asia-North  America, Mediterranean and Central Asia.       1.  There are altogether 4 endemic genera of Leguminosae in this region. Accord- ing to their morphological characters, systematic position and geographical distribution, it would appear that Salweenia and Piptanthus are Tertiary paleo-endemics, while Straceya and Cochlianths are neo-endemics. Salweenia and Piptanthus may be some of more primitive members in the subfamily Papilionasae and their allies are largely distributed in the southern Hemisphere.  The other two genera might have been derived from the northern temperate genus Hedysarum and the East Asian-North American genus Apios respectively, because of their morphological resemblance. They probably came into existanc during the uplifting of the Himalayas.       2. An analysis of temperate genera       There are twelve temperate genera of Leguminosae in the region, of which the more important elements in composition of flora, is Astragalus, Oxytropis and Cara- gana.       Astragalus  is a  cosmopolitan  genus comprising 2000 species, with its center distribution in Central Asia. 250 species, are from China so far known, in alpine zone of Southwest and Northwest, with 70 species extending farther to the Himalayas and Xizang Plateau.       Among them, there are 7 species (10%) common to Central Asia, 12 species (15.7%) to Southwest China and 40 species (60%) are endemic, it indicates that the differentia- tion of the species of the genus in the region is very active, especially in the subgenus Pogonophace with beards in stigma. 27 species amounting to 78.5% of the total species of the subgenus, are distributed in this region.  The species in the region mainly occur in alpine zone between altitude of 3500—300 m. above sea-level. They have developed into a member of representative of arid and cold alpine regions.      The endemic species of Astragalus in Xizang might be formed by specialization of the alien and native elements. It will be proved by a series of horizontal and vertical vicarism of endemic species.  For example, Astragalus bomiensis and A. englerianus are horizontal and vertical vicarism species, the former being distributed in southeast part of Xizang and the latter in Yunnan; also A. arnoldii and A. chomutovii, the former being an endemic on Xizang Plateau and latter in Central Asia.      The genus Oxytropis comprises 300 species which are mainly distributed in the north temperate zone. About 100 species are from China so far known, with 40 species extending to Himalayas and Xizang Plateau.  The distribution, formation and differ- entiation of the genus in this region are resembled to Astragalus.  These two genera are usually growing together, composing the main accompanying elements of alpine mea- dow and steppe.      Caragana is an endemic genus in Eurasian temperate zone and one of constructive elements of alpine bush-wood. About 100 species are from China, with 16 species in Xi- zang. According to the elements of composition, 4 species are common to Inner Mon- golia and Kausu, 4 species to Southwest of China, the others are endemic. This not only indicates that the species of Caragana in Xizang is closely related to those species of above mentioned regions, but the differentiation of the genus in the region is obviously effected by the uplifting of Himalayas, thus leading to the formations of endemic species reaching up to 50%.      3. An Analysis of Tropical Genera      There are 19 tropical genera in the region. They concentrate in southeast of Xizang and southern flank of the Himalayas. All of them but Indigofera and Desmodium are represented by a few species, especially the endemic species. Thus, it can be seen that they are less differentiated than the temperate genera.      However, the genus Desmodium which extends from tropical southeast and northeast Asia to Mexio is more active in differentiation than the other genera. According to Oha- Shi,s system about the genus in 1973, the species of Desmodium distributed in Sino-Hima- laya region mostly belong to the subgenus Dollinera and subgenus Podocarpium.  The subgenus Dollinera concentrates in both Sino-Himalaya region and Indo-China with 14 species, of which 7 species are endemic in Sino-Himalaya.  They are closely related to species of Indo-China, southern Yunnan and Assam and shows tha tthey have close con- nections in origin and that the former might be derived from the latter.      Another subgenus extending from subtropical to temperate zone is Podocarpium. Five out of the total eight species belonging to the subgenus are distributed in Sino- Himalaya and three of them are endemic.      An investigation on interspecific evolutionary relationship and geographic distribu- tion of the subgenus shows that the primary center of differentiation of Podocarpium is in the Sino-Himalaya region.      Finally, our survey shows that owing to the uplifting of the Himalayas which has brought about complicated geographic and climatic situations, the favorable conditions have been provided not only for the formation of the species but also for the genus in cer-tain degree.     

中国列当属的分类及与近缘属的关系

The morphological characters in the genus Orobanche were evaluated from the taxonomic point of view.  The author finds that the plants of this genus are relatively similar to each other in respect to characters of vegetative organs, fruits and seeds.  But the differences in the floral structures can be served as a basis for delimitating infrageneric taxa.   The seed coat of 18 species and pollen grains of  6 species were also examined under scanning electron microscope (SEM). They seem to have little significance for distinguishing species.       The result supports G. Beck’s (1930) division of the genus Orobanche into 4 sections, of which 2 occur in China, based on the characters of the inflorescence, bracteoles and calyx. The author considers that some characters, such as anther hairy or not, upper lip of corolla entire or not, lower lip longer or shorter than the upper one, the state of corolla-tube inflec-  tion and the hair type of filaments and plants, are important in distinguishing Chinese species.  A key to the species of Orobanche in China is given.       This genus consists of about 100 species, and is mostly confined to Eurasia, with over 60  species found in Caucasus and Middle Asia of USSR, where may be the mordern  distribu-  tional  centre.        Orobanche L. in China is represented by 23 species, 3 varieties and l forma. As shown in  Table 1, most species (12 species) are found in Xinjiang, which clearly shows a close floristic  relationship between this region and Middle Asia of USSR.  6 species are endemic to China,  of which 4 are confined to the Hengduan Mountains  (Yangtze-Mekong-Salwin divide).        The relationships between this genus and related ones of Orobanchaceae are also discussed.  The author holds the following opinions: the genus Phelypaea Desf. should be considered as a   member of Orobanche L. Sect. Gymnocaulis G. Beck,  the monotypic genus,   Necranthus A.   Gilli endemic to Turkey, is allied with Orobanche L. Sect.  Orobanche, the monotypic genus,   Platypholis Maxim, endemic to Bonin Is. of Japan, is far from Orobanche L. in relation and   should be regarded as a separate genus.        The 11 OTU’s, including all the sections of Orobanche L. and 7 genera of Orobanchaceae,   and 15 morphological characters were used in the  numerical  taxonomic treatment  to  test  the   above-mentioned  suggestions.   After standardization of characters, the correlation matrices were   computerized.  The correlation matrices were made to test the various clustering methods.   At    last the UPGMA clustering method was chosen and its result is shown in a phenogram.  The   result of numerical analysis is basically in accordance with the suggestions.     

星果草属、独叶草属、鸡爪草属的染色体观察和系统位置的探讨

1.  The present paper describes the observations of chromosome  numbers  and karyomorphology of 2 species of 2 endemic genera and I endemic species of Chinese Ranunculaceae: Asteropyrum peltatum (Franch.)  Drumm et Hutch. 2n=16, x=8; Kingdonia unifolia Balf. f. et W. W. Sm. 2n=18, x=9 and Calathodes oxycarpa Spra- gue 2n=16, x=8.  The chromosome counts of three ranunculaceous genera are repor- ted for the first time.       2.  The morphylogical, palynological and cytological date in relation to the syste- matic postition of Asteropyrum, Kingdonia and Calathodes within the family Ranun- culaceae are diseussed and resulted in following conclusions:       (1).  On the basis of the basic number x=8 in Asteropyrum, it is further con- firmed that this genus is distinct from the r elated genera such as Isopyrum, Dichocarp- um and other allied taxa.       The comparison of Asteropyrum with Coptis shows that they are identical in short chromosomes, with magnoflorina and benzylisaquinodine type of alkaloides, but dif- ferent from coptis in the chromosome numbers (T-type), pantocolpate pollens, united carpels and the dorsi-ventral type of petioles.  In view of these fundamental morpho- logical and cytological differences, Asterop yrum is better raised to the level of Tribe. However Asteropyrum and Coptis may represent two divaricate evolutional lines of Thalictroideae.       (2).  The systematic position of the genus Kingdonia has been much disputed in the past.  We support the view of Sinnote (1914), namely, the trilacunar in leaf traces “the ancient type”, appeared in the angiosperm line very early, while the uni- lacunar of Kingdonia may be derived from the trilacunar.  On the basis of the chromo- some numbers and morphylogical observation, the present writer accept Tamura’s and Wang’s treatment by keeping Kingdonia in Ranunculaceae instead of raising it to a family rank as has been been done by Forster  (1961).  Kingdonia and  Coptis are similar in having short chromosome with x=9, but with one-seeded fruits; therefore it is suggested that placed into Thalictroideae as an independent tribe, indicating its close relationship with Coptideae.        (3).  Comparing with its allies, Calathodes being with out petals, seems to be more  primitive than Trollius. But Calathodes differs from Trollius with R-type chromosomes in having T-type chromosome with x=8 and subterminal centromere.  Those charac- teristics show that it is very similar to the related genera of Thalictroideae.  But as Kurita already pointed out that most speci es of Ranunculus have usually large long chromosomes but some species have compar ativelly short chromosomes, therefore we regard T-type and R-type chromosomes appear independently in different subfamilies of Ranunculaceae. According to Tamura, G alathodes seems to be  closely related  to Megaleranthis, because of the resemblance in follicles.  But due to lack of cytological data of the latter genus, the relationship between the two genera still is not clear pen- ding further studies. From the fact that the  morphology and  chromosomes  of  the Calathodes differs from that of all other genera of the Helleboroideae, we consider Calathodes may form an independent tribe of its own with a closer relationship withTrollieae.     

木兰科分类系统的初步研究

A new system of classification of Magnoliaceae proposed.  This paper deals mainly with taxonomy and phytogeography of the family Magnoliaceae on the basis of external morphology, wood anatomy and palynology.  Different  authors have had different ideas about the delimitation of genera of this family, their controversy being carried on through more than one hundred years (Table I).  Since I have been engaged in the work of the Flora Reipublicae Popularis Sinicae, I have accumulated a considerable amount of information and material and have investigated the living plants at their natural localities, which enable me to find out the evolutionary tendencies and primitive morphological characters of various genera of the family.  According to the evolutionary tendencies of the characters and the geographical distribution of this family I propose a new system by dividing it into two subfamilies, Magnolioideae and Liriodendroideae Law (1979), two tribes, Magnolieae and Michelieae Law, four subtribes, Manglietiinae Law, Magnoliinae, Elmerrilliinae Law and Micheliinae, and fifteen genera (Fig. 1 ), a system which is different from those by J. D. Dandy (1964-1974) and the other authors.      The recent distribution and possible survival centre of Magnoliaceae. The members of Magnoliaceae are distributed chiefly in temperate and tropical zones of the Northern Hemisphere, ——Southeast Asia and southeast North America, but a few genera and species also occur in the Malay Archipelago and Brazil of the Southern Hemisphere. Forty species of 4 genera occur in America, among which one genus (Dugendiodendron) is endemic to the continent, while about 200 species of 14 genera occur in Southeast Asia, of which 12 genera are endemic.  In China there are about 110 species of 11 genera which mostly occur in Guangxi, Guangdong and Yunnan; 58 species and more than 9 genera occur in the mountainous districts of Yunnan.   Moreover,  one  genus (Manglietiastrum Law, 1979) and 19 species are endemic to this region.  The family in discussion is much limited to or interruptedly distributed in the mountainous regions of Guangxi, Guangdong and Yunnan.  The regions are found to have a great abundance of species, and the members of the relatively primitive taxa are also much more there than in the other regions of the world.      The major genera, Manglietia, Magnolia and Michelia, possess 160 out of a total of 240 species in the whole family.  Talauma has 40 species, while the other eleven genera each contain only 2 to 7 species, even with one monotypic genus.   These three major genera are sufficient for indicating the evolutionary tendency and geographical distribution of Magnoliaceae.  It is worthwhile discussing their morphological  characters  and distributional patterns as follows:      The members of Manglietia are all evergreen trees, with flowers terminal, anthers dehiscing introrsely, filaments very short and flat, ovules 4 or more per carpel.  This is considered as the most primitive genus in subtribe Manglietiinae.  Eighteen out of a total  of 35 species of the genus are distributed in the western, southwest to southeast Yunnan. Very primitive species, such as Manglietia hookeri, M. insignis  and M. mega- phylla, M. grandis, also occur in this region. They are distributed from Yunnan eastwards to Zhejiang and Fujian through central China, south China, with only one species (Manglietia microtricha) of the genus westwards to Xizang.  There are several species distributing southwards from northeast India to the Malay Archipelago (Fig. 7).      The members of Magnolia are evergreen and deciduous trees or shrubs, with flowers terminal, anthers dehiscing introrsely or laterally, ovules 2 per carpel, stipule adnate to the petiole.  The genus Magnolia is the most primitive in the subtribe Magnoliinae and is the largest genus of the family Magnoliaceae. Its deciduous species are distributed from Yunnan north-eastwards to Korea and Japan (Kurile N. 46’) through Central China, North China and westwards to Burma, the eastern Himalayas  and northeast India.  The evergreen species are distributed from northeast  Yunnan  (China)  to  the Malay Archipelago.  In China there are 23 species, of which 15 seem to be very primi- tive, e.g. Magnolia henryi, M. delavayi, M. officinalis and M. rostrata, which occur in Guangxi, Guangdong and Yunnan.      The members of Michelia are evergreen trees or shrubs, with flowers axillary, an- thers dehiscing laterally or sublaterally, gynoecium stipitate, carpels numerous or few. Michelia is considered to be the most primitive in the subtribe Micheliinae, and is to the second largest genus of the family.  About 23 out of a total of 50 species of this genus are very primitive, e.g. Michelia sphaerantha, M. lacei, M. champaca,  and  M. flavidiflora, which occur in Guangdong, Guangxi and Yunnan (the distributional center of the family under discussion)  and extend eastwards to Taiwan  of  China, southern Japan through central China, southwards to the Malay Archipelago through Indo-China. westwards to Xizang of China, and south-westwards to India and Sri Lanka (Fig. 7).      The members of Magnoliaceae are concentrated in Guangxi, Guangdong and Yunnan and radiate from there.  The farther away from the centre, the less members we are able to find, but the more advanced they are in morphology.  In this old geographical centre there are more primitive species, more  endemics  and  more monotypic genera. Thus it is reasonable to assume that the region of Guangxi, Guangdong and Yunnan, China, is not only the centre of recent distribution, but also the chief survival centreof Magnoliaceae in the world.     

知母染色体核型的研究

  A karyotypical analysis of Anemarrhena asphodeloides Bung. of the monotypic genus Anemarrhena Bung. (Liliaceae) was carried out for the first time. The number of chromo- somes in root-tip cell of the species was found to be 22, agreeing with that reported by Sato[12], although inconsistent in some other respects, such as position of  centromeres, length of chromosomes, and nucleoli, etc. (Table 1 ). According to the terminology defined by Levan et al.［8］, the karyotype formula is therefore 2n=22=2sm (SAT)+2sm+18m. Photomicrographs of the chromosome complements and idiogram of the karyotype are given   Fig. 1 and 2).       The karyotype of Anemarrhena asphodeloides shows explicitly to be asymmetrical, with three pairs of long chromosomes and eight pairs of short chromosomes. This specialized feature, when considered together with the rare occurrence of the basic chromosome number of 11 of the genus within the Tribe Asphodeleae of Liliaceae (see Table 1), suggests that the genus Anemarrhena is probably a rather specialized one, which has scarcely any intimate relationship with the other genera of the above tribe. The fact that this specialized karyotype is associated with certain trends of morphological specialization, such as flowers possessing three stamens only, gives support to the above suggestion. But, it is impossible to draw a more precise conclusion without a more thorough and comprehensive investigation of the species in question.     

蔷薇科太行花属系统位置的研究

 We have described a new genus Taihangia, collected from, the south part of Taihang Mountain in northern China. At the same time, comparative studies on Taihangia with its related genera have been made in various fields including external morphology, anatomy of carpels, chromosome and pollen morphology by light, scanning and transmission electron microscope. In addition, isoperoxidases of two varietier were analysed by means of polya-crylamide gel slab electrophoresis. The preliminary results are as follows:       Morphology: The genus Taihangia is perennial and has simple leaves, occasionally with 1—2 very small reduced lobes on the upper part of petiole; flowers white, andromo- noecious and androdioecious, terminal, single or rarely 2 on a leafless scape; calyx and cpicalyx with 5 segments; petals 5; stamens numerous; pistils numerous, with pubescent styles, spirally inserted on the receptacle in bisexual flowers, but with less number of abortive and glabrous pistils in male flowers.       In comparison with the related genera such as Dryas, Geum, Coluria and Waldsteinia, the new genus has unisexual flowers and always herbaceous habit indicating its advanced feature but the genus has a primitive style with thin and short hairs as compared with the genus Dryas which has long, pinnately haired styles, a character greatly facilitamg anemo-choric dissemination. The styles of Taihangia are slender and differ from those of the ge-nus Geum which are articulate, with a persistent hooked rostrum, thus adapting to epizo-ochoric dissemination to a higher degree.       The anatomy of carpels shows the baral position of ovules in the genus Taihangia like those in other related genera such as Dryas, Geum, Acomastylis, Coluria and Waldsteinia. This suggests that the new genus and its related ones are in a common evolutionary line as compared with the other tribes which have a pendulous ovule and represent a separate evolutionary line in Rosaceae. Dorsal and ventral bundles in carpels through sections are free at the base. Neither fusion, nor reduction of dorsals and vertrals. are observed. This shows that the genus Taihangia is rather primitive.       Somatic chromosome: All the living plants, collected from both Honan and Hopei Provinces were examined. The results show that in these plants the chromosome number is 2n= 14, and thus the basic number of chromosome is x=7. Such a diploid genus is first found in both anemochoric and epizoochoric genera. Therefore, in this respect Taihangia is primitive as compared with herbaceous polyploid genus Geum and related ones.      Pollen: The stereostructure shown by scanning electron microscope reveals  that  the pollen grains of the genus Taihangia are ellipsoid and 3-colporate. There are two types of exine sculpture. One is rather shortly striate and it seems rugulate over the pollen surface; the other is long-striate. The genus Dryas differs in having only short and thick striae over the surface. The genus is similar to the genera Geum, Coluria and Waldsteinia in colpustype, but differs from them in that they all have long, parallel striae which are distributed along the meridional line.       In addition, under transmission electron microscope, the exine in the Taihangia and related genera Acomastylis, Geum, Coluria, Waldsteinia and Dryas has been shown to be typically differentiated into two distinct layers, nexine and sexine. The nexine, weakly statined, appears to consist of endoxine with no foot-layer, in which the columellae are fused, and which is thicker beneath the apertures. The sexine is 2-layered, consisting of columellae and tectum. Three patterns of tectum can be distinguished in the tribe Dryadeae: the first, in the genera Taihangia, Acomastylis, Geum, Coluria and Waldsteinia, is tectate-imperforate, with the sculpturing elements both acute and obtuse at the top and broad at the base; the second, in the genus Dryas, is semitectate, with the sculpturing elements shown in ultrathin sections rod-like and broader at the top than at the base or as broad at the top as at the base, and the third, tectate-perforate, with the sculpturing elements different in size. From the above results, the herbaceous groups and woody ones  have palynologically evolved in two distinct directions, and the genus Taihangia is related to other herbaceous genera such as Acomastylis, Geum, Coluria and Waldsteinia, as shown in the electron microphotographs of ultrathin sections. The genus Taihangia, however, is different from related herbaceous genera in that the pollen of Taihangia is dimorphic, i.e. in addition to the above pattern of pollen another one of the exine in Taihangia is rugulate, with the sculpturing elements shown in the ultrathin sections being obtuse or emarginate and nearly as broad at the top as at the base.      The interesting results obtained from the comparative analysis of morphology, ana- tomy of carpels, chromosome countings, microscopic and submicrosocopic structures of pollen may enable us to evaluate the systematic position of Taihangia and to throw a new light on evolution of the tribe Dryadeae. It is well known that the modes of dissemination of rosaceous fruits play an important role in the expansion and evolution of the family. The follicle is the most primitive and the plants with follicles, like the Spiraeoideae, are mostly woody and mesic, while the achene, drupe and pyrenarium are derived. In Rosoideae  having a achene is a common feature. Particularly in the tribe Dryadeae, which is distinguished from the other related tribes by having orthotropous ovules, the methods of dissemination of fruits have developed in three distinct specialized directions: anemochory with long, plumose styles (e.g. Dryas), formicochory or dispersed by ants or other insects, with the deciduous styles (e.g. Waldsteinia and Collria),and epizoochory with the upper deciduous stigmatic part and the lower persistent hooked rostrum, an  adhesive organ favouring  epizoochory dissemination (e. g. Geum and related taxa). Taihangia is a genus endemic to mesophytic forest area of northern China. Due to its narrow range and specific habit as well as pubescent styles, neither perfectly adapted to anemochory nor to epizoochory, the genus  Taihangia might be a direct progeny of the ancestry of anemochory. Maintaining the diploidy and having an ntermediate sculptural type of pollen, the new genus might probably represent a linkage between anemochory and zoochory (including epizoochory and dispersed by ants).       Experimental evidence from isoperoxidases shows the stable zymograms of root and roostoks. The anodal isozyme of T. rupestris var. rupestris may be divided into 6 bands: A, B, C, D, E, F, and T. rupestris var. ciliata into 4 bands: A, B, C, G. The two varietiesof the species share 3 bands: A, B, C. However, D, E and F bands are characteristic of var. rupestris and G band is limited to var. ciliata. As far as the available materials are concerned, the analysis of isoperoxidases supports the subdivision of the species into two varieties.     

两种蕨类植物的新命名

 The fern genus Diplaziopsis C. Chr. of Index Filicum has long been considered as a monotypic one, with the sole species, D. javanica (B1.) C. Chr. from tropical Asia.  In 1906, H. Christ described a second species, Allantodia cavaleriana Christ (=D. cavale- riana C. Chr.) from Kweichow, West China, but this was since not fully recognized by fern students in general, being often considered as a variety of the first species.  This is certainly a mistake, as is shown by ample herbarium specimens today.  In the recent work on the genus, the writer has found among the herbarium material two additional new species from China, thus bringing the genus up to four species in Asia, mainly from China, where, as it is, the genus has its center of development from the long past.      Phylogenetically, Diplaziopsis C. Chr. represents one of the offshoots from the great stock of diplazioid ferns, of which the genus Diplazium Sw. constitutes the main body of the group and from which our genus differs chiefly in its leaves of a thin texture with reticulated venation, but not so much in its type of indusium as it has generally been emphasized by most botanists in the past, for, as it is, the type of indusium in Di- plaziopsis also prevails in many species of Diplazium, for which C. B. Clarke (Trans. Linn. Soc. ser. 2, Bot. I:495, 1880) created, but really superfluously, a subgenus Pseudal- lantodia, about which the writer will dwell in another paper in the near future.  Suffice it to say here that the indusium in Diplaziopsis as revealed by the species treated here is, indeed, typical of diplazioid ferns, only often, as it happens, with  its  adaxial  edge pressed so tight under the expanding sorus that it is unable to open freely along its upper free edge and, as a result, its thin vaulted back bursts open from the pressure of the ex- panding sorus underneath.      As a result of the present study, following four species of the genus have been re- cognized.      Diplaziopsis javanica (B1.) C. Chr. Ind. Fil. (1905) 227.      Wide spread in tropical Asia, northwardly to Bakbo and the southern part of Yun- nan, China.      D. cavaleriana (Christ) C. Chr. Ind. Fil. Suppl. I (1913) 25.      Ranges from West China through northern part of Fukien of East China to Japan.      D. intermedia Ching, sp. nov.      Endemic in West China:  Mt. Omei, Szechuan, and Kweichow.      D. hainanensis Ching, sp. nov.      In conclusion, it may be pointed out that with the modern plant taxonomy pursued in a more efficient manner than in the past, and especially by the introduction of the cytotaxonomic methods, the so-called “monotypic genera”, as conceived by the orthodox systematists, will continue to prove, to a great extent, to be lack of  enough  scientific ground.  The fact that the “monotypic genus” of Diplaziopsis C. Chr. is now found to be a genus of four well-defined species is once again an instance to illustrate the pointat issue.     

浙江省薯蓣科植物的数量分类研究

以浙江省薯蓣科全部植物——薯蓣属17种和1变种为18个分类单位，以形态为主兼顾组织结构、化学成分及地理分布，列出72项性状，对这些植物进行了数量分类。用最短距离法、最长距离法、中间距离法甲、中间距离法乙、重心法、类平均法以及离差平方和法的系统聚 类，探索了这些植物彼此间的亲缘关系，认为以前对它们的经典分类处理基本合理。由于”薯莨”性状明显特殊，建议成立新组——薯莨组。     

柠檬酸溶胶凝胶法制备铈锆固溶体的物性和氧化还原特性研究

以柠檬酸作为胶凝剂，采用溶胶凝胶法制备了一系列CeXZr1-XO2复合氧化物，分别经600℃和900℃焙烧，应用XRD检测，发现均已形成铈锆固溶体。并应用BET、XRD和TPR对其表相、晶相结构和氧化还原性质进行了分析，发现随着固溶体中铈比例的增加，固溶体稳定性降低；其储氧量测定显示铈锆固溶体用作三效催化剂助剂的最佳比例为1：1。经900℃高温焙烧以后，固溶体晶粒增大不明显，表明本方法适合制备高比表面积和高储氧量的铈锆固溶体；TPR还原峰温度降低，推测可能在于柠檬酸络合物完全分解所致。     

反射仪—K+试纸法快速测定烟草含K+水平的方法研究

比较了反射仪—K+试纸法(K+ts-fl)、原子吸收光谱法、ICP–aeS法测定烟株叶脉汁液中的K+含量的三种方法，确定反射仪—K+试纸法测定烟株钾含量的可行性及其最佳测定范围；在烤烟不同生育期，应用反射仪—K+试纸法对烟株叶片不同部位含K+水平的测定，确定其最佳测定时期及部位。研究结果表明，反射仪—K+试纸法与原子吸收光谱法、ICP–aeS法测定结果差异不显著，应用反射仪—K+试纸法进行烟株钾素快速诊断可行，且具有时间短，简单快速，易操作等优点。反射仪测定K+的稳定线性范围是0.30g/L~0.9g/L；在不同时期对烟株不同部位测定研究表明，最佳诊断部位为烟株第二平展叶叶脉基部2cm段。     

论胡桃科植物的地理分布

 The present paper aims to discuss the geog raphical distribution of the Juglandaceae on the basis of unity of the phylogeny and the process of dispersal in the plants.       The paper is divided into the following three parts:       1.  The systematic positions and the distribution patterns of nine living genera in the family Juglandaceae (namely, Engelhardia, Oreomunnea, Alfaroa, Pterocarya, Cyclo- carya, Juglans, Carya, Annamocarya and Platycarya) are briefly discussed.  The evolu- tional relationships between the different genera of the Juglandaceae are elucidated. The fossil distribution and the geological date of the plant groups are reviewed.  Through the analysis for the geographical distribution of the Juglandaceous genera, the distribu- tion patterns may be divided as follows:       A.  The tropical distribution pattern       a. The genera of tropical Asia distribution: Engelhardia, Annamocarya.       b. The genera of tropical Central America distribution: Oreomunnea, Alfaroa.       B.  The temperate distribution pattern       c. The genus of disjunct distribution between Western Asia and Eastern Asia: Pterocarya.       d. The genus of disjunct distribution between Eurasia and America: Juglans.      e. The genus of disjunct distribution between Eastern Asia and North America: Carya.      f. The genera whose distribution is confined to Eastern Asia: Cyclocarya, Platy- carya.      2.  The distribution of species      According to Takhtajan’s view point of phytochoria, the number of species in every region are counted.  It has shown clearily that the Eastern Asian Region and the Coti- nental South-east Asian Region are most abundant in number of genera and species. Of the 71 living species, 53 are regional endemic elements, namely 74.6% of the total species. The author is of the opinion that most endemic species in Eurasia are of old endemic nature and in America of new endimic nature.  There are now 7 genera and 28 species in China, whose south-western and central parts are most abundant in species, with Pro- vince Yunnan being richest in genera and species.      3.  Discussions of the distribution patterns of the Juglandaceae A.       The centre of floristic region B.         The centre of floristic regions is determined by the following two principles:  a. A large number of species concentrate in a district, namely the centre of the majority; b.  Species of a district can reflect the main stages of the systematic evolution of the Juglandaceae, namely the centre of diversity.  It has shown clearly that the southern part of Eastern Asian region and the northern part of Continental South-east Asian Region (i.c. Southern China and Northern Indo-China) are the main distribution centre of the Juglandaceae, while the southern part of Sonora Region and Caribbean Region  (i.c. South-western U.S.A., Mexico and Central America) are the secondary distribution centre.       As far as fossil records goes, it has shown that in Tertiary period the Juglanda- ceae were widely distributed in northern Eurasia and North America, growing not only in Europe and the Caucasus but also as far as in Greenland and Alaska.  It may be considered that the Juglandaceae might be originated from Laurasia.  According to the analysis of distribution pattern for living primitive genus, for example, Engelhar- dia, South-western China and Northern Indo-China may be the birthplace of the most primitive Juglandaceous plants.  It also can be seen that the primitive genera and the primitive sections of every genus in the Juglandaceae have mostly distributed in the tropics or subtropics. At the same time, according to the analysis of morphological cha- racters, such as naked buds in the primitive taxa of this family, it is considered that this character has relationship with the living conditions of their ancestors.  All the evidence seems to show that the Juglandaceae are of forest origin in the tropical moun- tains having seasonal drying period.       B.  The time of the origin       The geological times of fossil records are analyzed. It is concluded that the origin of the Juglandaceae dates back at least as early as the Cretaceous period.       C.  The routes of despersal       After the emergence of the Juglandaceous plant on earth, it had first developed and dispersed in Southern China and Indo-China.  Under conditions of the stable tempera- ture and humidity in North Hemisphere during the period of its origin and development, the Juglandaceous plants had rapidly developed and distributed in Eurasia and dis- persed to North America by two routes: Europe-Greenland-North  America  route  and Asia-Bering Land-bridge-North America route.  From Central America it later reached South America.      D.  The formaation of the modern distribution pattern and reasons for this forma- tion.      According to the fossil records, the formation of two disjunct areas was not due to the origin of synchronous development, nor to the parallel evolution in the two con- tinents of Eurasia and America, nor can it be interpreted as due to result of transmis- sive function.  The modern distribution pattern has developed as a result of the tectonic movement and of the climatic change after the Tertiary period.  Because of the con- tinental drift, the Eurasian Continent was separated from the North American Conti- nent, it had formed a disjunction between Eurasia and North America. Especially, under the glaciation during the Late Tertiary and Quaternary Periods, the continents in Eu- rasia and North America were covered by ice sheet with the exception of “plant refuges”, most plants in the area were destroyed, but the southern part of Eastern Asia remained practically intact and most of the plants including the Juglandaceae were preserved from destruction by ice and thence became a main centre of survival in the North Hemisphere, likewise, there is another centre of survival in the same latitude in North America and Central America.      E.  Finally, the probable evolutionary relationships of the genera of the Juglanda-ceae is presented by the dendrogram in the text.     

披碱草属与大麦属系统关系的研究

禾本科中，披碱草属Elymus L.为多倍体属，约含150余种；大麦属Hordeum L.具二倍体和多倍体，约有40余种，该两属均广泛地分布于全球温带地区。该两属，尤其是披碱草属的系统分类较为困难。基于形态学的传统研究认为这两个属的系统关系较远，而细胞学研究的资料却表明，披碱草属的H染色体组起源于大麦属。笔者对来源不同的披碱草属和大麦属的物种进行了远缘杂交，并对其属间杂种F1的减数分裂中期I染色体配对行为进行了分析。结果表明，若以披碱草属作母本，该两属有相对较高的杂交亲合力，通过对杂种幼胚进行分割和离体培养，也能获得杂种F1植株。属间杂种植株的形态介于双亲之间，但更接近于披碱草属，杂种的生殖器官发育不健全，而且所有的杂种F1均完全不育。细胞学的观察结果表明，这两个属间的杂种F1通常具有较低的减数分裂中期I染色体配对数，但有较大的变异。通过笔者的工作及掌握的形态学和细胞学的资料分析认为：披碱草属和大麦属的亲缘关系较为复杂，不能一概而论。含H染色体组的披碱草属和大麦属物种有着较近的亲缘关系，但这两个属中所含的H染色体组已产生了程度不同的分化；不含H染色体组的披碱草属及大麦属的物种具有较远的亲缘关系。     

沙参属10个种的染色体研究

 本文报道了我国黑龙江产桔梗科沙参属的10种1变种的染色体数目和核型，对其中     7种作了减数分裂行为的观察。  其中6种1变种为首次报道，并发现2n=68的4x种。该     属染色体基数多为17(x＝17)，但Adenophora trachelioides和A．remotiflora的基数为18     (x=18)，为该属独特基数。核型的共同特征是：小型，以中部(m)、近中部(sm)着丝点     染色体为主，至少具一对近端着丝点染色体和一对随体染色体。该属染色体的演化处于二种     水平：  数目变化(包括多倍化和非整倍体变化)和结构变异。  多倍化是该属物种形成的主要     途径之一。结合其它性状讨论了这些种的分类，并确立1个四倍体新种(A. amurica)和1个新组合(A．pereskiifolia ssp．alternifolia)。     

裂瓜属的研究

The classical and numerical taxonomy, palynology and the geographical dis- tribution of the Genus Schizopepon are dealt with in the present paper.  Having comme- nted on various opinions regarding the systematic position of the genus, the present au- thors consider that C. Jeffrey’s treatment of Schizopepon as a new and monogeneric tri- be, Schizopeponeae, should be supported.      The gross morphological characters in the genus are assessed from the taxonomic point of view.  Some characters, such as stamens with an elongated connective or not, different insertions of ovules and various forms of ovaries and fruits, may be used for distinguishing subgenera.      The pollen grains of all the species were observed under light microscope (LM) and scanning electron microscope (SEM).  The results show that a strong differentiation has taken place in the pollen of the genus, and in consequence it may be regarded as an important basis for dividing subgenera and species. Especially it should be pointed out that degrees of development of colpi and positions of ora are positively correlated with the external characters used for distinguishing subgenera.      According to the morphological and palynological characters, the genus Schizopepon may be divided into three subgenera and eight species: 1. Subgenus Schizopepon: 5 spe- cies, S. bryoniaefolius Maxim., S. monoicus A. M. Lu et Z. Y. Zhang, S. dioicus Cogn., S. longipes Gagnep. and S. macranthus Hand.-Mazz.; 2. Subgenus Rhynchocarpos A. M. Lu et Z. Y. Zhang: 1 species, S. bomiensis A. M. Lu et Z. Y. Zhang; 3. Subgenus Neoschi- zopepon A. M. Lu et Z. Y. Zhang: 2 species, S. bicirrhosus (C. B. Clarke) C. Jeffrey and S. xizangensis A. M. Lu et Z. Y. Zhang.      The 8 OTU’s including all the species of this genus and 31 characters, of which 16 are morphological characters and 15 palynological characters, were used in the numerical taxonomic treatment.  After standardization of characters, the correlation and distance matrices were computed.  The correlation matrices are made to test the various clustering methods.  At last, the UPGMA clustering method was selected and its result is shown in the form of phenogram.  The result of numerical analysis is similar to that of the classical classification.      Schizopepon Maxim. is a genus of East Asia-Himalayan distribution. China has all 8 species and 2 varieties, of which 6 species are endemic. Based on the statistics of spedies number, the distribution centre of the genus is considered to be in the Hengduan Mountains (Yangtze-Mekong-Salwin water divides) and the adjacent areas of the southwest China.