中国凤尾蕨属细胞学的初步研究

本文报道了中国产10种凤尾蕨属植物和2种蕨属和栗蕨属植物的细胞学研究结果。  在凤尾蕨   属中，8种是多倍体或具有多倍体细胞型，4种是无性孢子繁殖的后代，6种实际上是种复合体或种复合   体成员，单纯有性生殖的二倍体只有2种。凤尾蕨至少具有9条孢子发生路线，在其同一个体上除了产   生二倍孢子外，还可能产生少量加倍或多倍孢子；广义的蜈蚣草实际上是个种复合体，其祖先的二倍体   细胞型广泛分布于中国亚热带地区说明这里可能是该复合体的起源中心；岩凤尾蕨具有不寻常的染色  体数目n＝55，这说明在本属和本科中可能存在着非整倍体进化。     

蜘蛛抱蛋属(百合科)的细胞分类学研究(1)——四川七个种的核型

In the work mitotic chromosomes in root-tips of 7 species native to Sichuan Pro- vince were examined and their karyotypes were analysed based on 7-8 cells at mitotic meta- phase, using Levan et al.'s (1964) nomenclature. The list of species and origin of the materials used in this work are provided in the appendix.  The micrographs of mitotic metaphase of all the materials are shown in Plates 1 and 2; the idiograms in Fig. 1, 1-9, and the parameters of chromosomes are provided in Tables 1-9.  All the chromosome countings and karyotypes in this paper are reported for the first time.      Characteristics of the karyotypes may be summarized as follows:      1.  2n=38 are found in all the materials except A. sichuanensis, which has 3 cytotypes, i.e, 2n=38+5B, 2n=38+lB and 2n=38+OB (Plate 2, 1-2, Fig. 1, 5-6).      2.  The karyotypes are of two major types: the karyotype of A. flaviflora falls into 3C in Stebbins's (1958, 197l) classification of karyotypes and those of all the other species into 2C. The two types are also different from each other in the number of large and medium-sized chromosomes pairs and morphology of the first pair of chromosomes (compare Plate 2, 5, Fig. 1, 9 with the other micrographs and idiograms).A. flaviflora with the karyotype 3C also differs from the other species in a series of gross morphological characters: the species is of a yellow and campanulate corolla.      3.  The species with caespitose leaves (A. caespitosa and A. omeiensis) have essentially the same karyotype, which is rather different from those of the species with scattered leaves. There are two pairs of small sm chromosomes (arm ratio ≥1.90) in the former karyotype (Tables 1 and 3), but all the small chromosomes are m or sm with arm ratio <1.80 in the latter karyotypes.     

变异铁角蕨和其混淆种的正误研究

The present paper aims at clarifying the identity of Asplenium varians Wall. and its related species for the forthcoming Flora of China vol. 4. Wide-spread in the northern, north-western and south-western parts of China and adjacent regions and growing in exposed rock crevices is the group of ferns in question, which has hi- therto been taxonomically confused in the botanical literature.  Many distinct and related species were previously identified as Aspl. varians, a most wide-spread fern with very variable fronds which may vary from 10 to 30 cm in height and from simply pinnate to fully bipinnate in the degree of pinnation in a single clump under different habitats.     

沙参属10个种的染色体研究

 本文报道了我国黑龙江产桔梗科沙参属的10种1变种的染色体数目和核型，对其中     7种作了减数分裂行为的观察。  其中6种1变种为首次报道，并发现2n=68的4x种。该     属染色体基数多为17(x＝17)，但Adenophora trachelioides和A．remotiflora的基数为18     (x=18)，为该属独特基数。核型的共同特征是：小型，以中部(m)、近中部(sm)着丝点     染色体为主，至少具一对近端着丝点染色体和一对随体染色体。该属染色体的演化处于二种     水平：  数目变化(包括多倍化和非整倍体变化)和结构变异。  多倍化是该属物种形成的主要     途径之一。结合其它性状讨论了这些种的分类，并确立1个四倍体新种(A. amurica)和1个新组合(A．pereskiifolia ssp．alternifolia)。     

毛茛科金莲花族和升麻族细胞学的比较研究

对毛茛科金莲花族Trolliese和升麻族Cimicifugeae的细胞学进行了比较研究。发现驴蹄草Caltha palustris L．在云南西北部形成一个多倍体系列(2n=32，48，64)，四倍体细胞型(2n=4x=32)较为常见，其核型有明显的居群间变异。驴蹄草属Caltha、鸡爪草属Calathodes、Megaleranthis以及金莲花属Trollius的染色体在大小上基本相似，都属于中等大小的R-型染色体。细胞学和花粉学证据都支持鸡爪草属与Megaleranthis和金莲花属有较近的亲缘关系。铁破锣属Beesia、Anemonopsis、黄三七属Souliea、升麻属Cimicifuga以及类叶升麻属Actaea的核型彼此基本相似，在染色体大小和形态上都与驴蹄草属、鸡瓜草属、Megaleranthis以及金莲花属的核型明显有别。细胞学证据表明铁破锣属应是升麻族中的成员。     

西藏高原植物区系中特有属的研究

The Xizang (Tibetan) flora with numerous endemics is of importance in Chi- nese flora.   According to recent statistics there are in Xizang 27 genera of  spermatophytes endemic to China, being only 2.25% percent of the total number of genera in the Xizang flora. Four of them are regarded as palaeoendemics (14.81%) and the others as neoendemics (85.19%). These endemic genera, of 30 species and 3 varieties, belong to 17 families, of which, Umbelli- ferae contains 6 genera, 7 species and 3 varieties; Compositae has 6 genera and 7 species, and Gentianaceae 1 genus and 2 species.  All the other families each comprises one genus with a single species.       The cosmopolitan families together comprising 14 genera with 15 species have the highest perecentage (52.92%) and the tropical ones (5 families, 5 genera with 5 species) come to the next (29.42%), followed by the temperate ones (3 families, 10 genera with 10 species) (17.66%). It shows that these endemic genera are obviously related to the tropical flora and temperate one in essence.        According to the number of species, the genera endemic to China and occurring in Xi- zang flora may be grouped as fallows. Monotypic endemic ones 14 (51.85%) Ditypic endemic ones 6 (22.22%) Oligotypic endemic ones 4 (14.81%) Small endemic ones 3 (11.11%)        The formation of the endemic genera is correlated with the topography, climate and en- vironmental conditions, and they may have resulted from the diversification in geography and climatic influence for a long time.  The southeastern part of Xizang Plateau is of very diverse ecological conditions, with the adequate precipitation, which may explain the concentration of these endemic genera in this region.        The largest similarity coefficient (38.30%) of the genera endemic to China and occurring in Xizang is with those in Qinghai Plateau, next, with those in Yunnan and in Sichuan pro- vinces (both 27.60%), which shows that these endemic genera are related to the floras of the regions mentioned above.        The difference in the horizontal distribution of these endemic genera is obviously between the southern and northern parts of Xizang Plateau.  The vertical distribution of the genera is also rather obvious, from 800 m to 5200 m above sea level, but concentrated in the zone of 3000 m to 4500 mm.  Therefore their occurrence in Xizang is not only affected by the historical environmental conditions but also controlled by the horizontal and vertical distribution.      The origin and evolution of some endemic genera, such as Psammosilene, Parateropyrum, Sphaerotylos, Salweenia, Ajaniopsis, Xizangia, Sinoleontopodium, are discussed in this paper.      Parateropyrum, a monotypic palaeotropic endemic, belongs to the tribe Atraphaxideae in- cluding Atraphaxis, Calligonum and Pteropyrum.  It may be a comparatively advanced group in the tribe, and is closely related to the genus Pteropyrum  which is  distributed in western Asia.  The genus Parapteropyrum has possibly survived as a palaetropic-tertiary  relic in this region.      Sphaerotylos, a member of the subtribe Sphaerotylinae, the tribe Boehmerieae in the family Urticaceae, is a comparatively primitive genus in the tribe Boehmerieae so far known.  As the other subtribes, such as Boehmerinae, Sarconchlamydinae, Orecnidinae and Maoutinae, are dis- tributed in the tropics, rarely in the subtropics, the genus is no doubt a palaetropic -tertiary relic.      Sinoleontopodium, belonging to the tribe lnuleae in Compositae, is also related to the ge- nus Leontopodium.  It is probable that the genus Sinoleontopodium arised later than the other.       We come to the conclusion that the southern part of Xizang Plateau is also one of thecentres of the origin and differentiation of genera endemic to China.     

薄鳞蕨属(中国蕨科)植物的孢子形态研究

对薄鳞蕨属5种3变种的孢子形态进行了光镜和扫描电镜观察。该属植物的孢子明显地分为两种类型：薄叶薄鳞蕨和杜氏薄鳞蕨的孢子为近球形，周壁为鸡冠状或拟网状纹饰；而华北薄鳞蕨、华西薄鳞蕨和绒毛薄鳞蕨的孢子为四面体状球形，周壁表面近光滑。第一种类型的孢子也存在于粉背蕨属，因此薄鳞蕨属和粉背蕨属的关系仍需进一步的研究。