西藏豆科植物区系的形成与分化

 Xizang (Tibet) is rich in Leguminosae flora, comprising 41 genera and 254 species so far known, exclusive of the commonly cultivated taxa (including 11 genera and 16 species). There are 4 endemic genera (with 8 species), 10 temperate genera (with 175 species) and 19 tropical genera (with 46 species) as well as the representatives of those genera whose distribution centers are in East Asia-North  America, Mediterranean and Central Asia.       1.  There are altogether 4 endemic genera of Leguminosae in this region. Accord- ing to their morphological characters, systematic position and geographical distribution, it would appear that Salweenia and Piptanthus are Tertiary paleo-endemics, while Straceya and Cochlianths are neo-endemics. Salweenia and Piptanthus may be some of more primitive members in the subfamily Papilionasae and their allies are largely distributed in the southern Hemisphere.  The other two genera might have been derived from the northern temperate genus Hedysarum and the East Asian-North American genus Apios respectively, because of their morphological resemblance. They probably came into existanc during the uplifting of the Himalayas.       2. An analysis of temperate genera       There are twelve temperate genera of Leguminosae in the region, of which the more important elements in composition of flora, is Astragalus, Oxytropis and Cara- gana.       Astragalus  is a  cosmopolitan  genus comprising 2000 species, with its center distribution in Central Asia. 250 species, are from China so far known, in alpine zone of Southwest and Northwest, with 70 species extending farther to the Himalayas and Xizang Plateau.       Among them, there are 7 species (10%) common to Central Asia, 12 species (15.7%) to Southwest China and 40 species (60%) are endemic, it indicates that the differentia- tion of the species of the genus in the region is very active, especially in the subgenus Pogonophace with beards in stigma. 27 species amounting to 78.5% of the total species of the subgenus, are distributed in this region.  The species in the region mainly occur in alpine zone between altitude of 3500—300 m. above sea-level. They have developed into a member of representative of arid and cold alpine regions.      The endemic species of Astragalus in Xizang might be formed by specialization of the alien and native elements. It will be proved by a series of horizontal and vertical vicarism of endemic species.  For example, Astragalus bomiensis and A. englerianus are horizontal and vertical vicarism species, the former being distributed in southeast part of Xizang and the latter in Yunnan; also A. arnoldii and A. chomutovii, the former being an endemic on Xizang Plateau and latter in Central Asia.      The genus Oxytropis comprises 300 species which are mainly distributed in the north temperate zone. About 100 species are from China so far known, with 40 species extending to Himalayas and Xizang Plateau.  The distribution, formation and differ- entiation of the genus in this region are resembled to Astragalus.  These two genera are usually growing together, composing the main accompanying elements of alpine mea- dow and steppe.      Caragana is an endemic genus in Eurasian temperate zone and one of constructive elements of alpine bush-wood. About 100 species are from China, with 16 species in Xi- zang. According to the elements of composition, 4 species are common to Inner Mon- golia and Kausu, 4 species to Southwest of China, the others are endemic. This not only indicates that the species of Caragana in Xizang is closely related to those species of above mentioned regions, but the differentiation of the genus in the region is obviously effected by the uplifting of Himalayas, thus leading to the formations of endemic species reaching up to 50%.      3. An Analysis of Tropical Genera      There are 19 tropical genera in the region. They concentrate in southeast of Xizang and southern flank of the Himalayas. All of them but Indigofera and Desmodium are represented by a few species, especially the endemic species. Thus, it can be seen that they are less differentiated than the temperate genera.      However, the genus Desmodium which extends from tropical southeast and northeast Asia to Mexio is more active in differentiation than the other genera. According to Oha- Shi,s system about the genus in 1973, the species of Desmodium distributed in Sino-Hima- laya region mostly belong to the subgenus Dollinera and subgenus Podocarpium.  The subgenus Dollinera concentrates in both Sino-Himalaya region and Indo-China with 14 species, of which 7 species are endemic in Sino-Himalaya.  They are closely related to species of Indo-China, southern Yunnan and Assam and shows tha tthey have close con- nections in origin and that the former might be derived from the latter.      Another subgenus extending from subtropical to temperate zone is Podocarpium. Five out of the total eight species belonging to the subgenus are distributed in Sino- Himalaya and three of them are endemic.      An investigation on interspecific evolutionary relationship and geographic distribu- tion of the subgenus shows that the primary center of differentiation of Podocarpium is in the Sino-Himalaya region.      Finally, our survey shows that owing to the uplifting of the Himalayas which has brought about complicated geographic and climatic situations, the favorable conditions have been provided not only for the formation of the species but also for the genus in cer-tain degree.     

西藏菊科植物

 1)  The Compositae in Tibet so far known comprise 508 species and 88 genera, which nearly amounts to one fourth of the total number of genera and one third of the total number of species of Compositae in all China, if the number of 2290 species and 220 genera have respectively been counted in all China. In Tibet there are all tribes of Com- positae known in China, and surprisingly, the large tribes in Tibetan Compositae are also large ones in all China and the small tribes in Tibet are also small ones in all China. Generally speaking, the large genera in Tibet are also large ones in all China and the small genera in Tibet are likewise small ones in all China. In this sense it is reasonable to say that the Compositae flora of Tibet is an epitome of the Compositae flora of all China.      In the Compositae flora of Tibet, there are only 5 large genera each containing 30 species or more. They are Aster, Artemisia, Senecio, Saussurea and Cremanthodium. And 5 genera each containing 10—29 species. They are Erigeron, Anaphalis, Leontopodium, Ajania, Ligularia and Taraxacum. In addition, there are 77 small genera, namely 87% of the total of Compositae genera in Tibet, each comprising 1—9 species, such as Aja-niopsis, Cavea and Vernonia, etc.      2)  The constituents of Compositae flora in Tibet is very closely related to those of Sichuan-Yunnan provinces with 59 genera and 250 species in common. Such a situation is evidently brought about by the geographycal proximity in which the Hengtuang Shan Range links southeastern and eastern Tibet with northern and northwestern Sichuan- Ynnnan.  With India the Tibetan Compositae have 59 genera and 132 species in common, also showing close floristic relationships between the two regions. Apparently the floris- tic exchange of Compositae between Tibet and India is realized by way of the mountain range of the Himalayas.  The mountain range of the Himalayas, including the parallel ranges, plays a important role as a bridge hereby some members of the Compositae of western or northern Central Asia and of the northern Africa or of western Asia have migrated eastwards or southeastwards as far as the southern part of Fibet and northern part of India, or hereby some Compositae plants of eastern and southeastern Asia or Asia Media have migrated northwestwards as the northern part of Central Asia.      Some of the species and genera in common to both Tibet and Sinjiang indicate that this weak floristical relationship between these regions is principally realized through two migration routes: one migration route is by way of the Himalayas including the parallel ranges to Pamir Plataeu and Tien Shan, or vice versa. The other migration route is by way of northern Sinjiang to Mongolia, eastern Inner Mongolia, southwards to Gansu, Qinghai (or western Sichuan), eastern Tibet up to the Himalayas, or vice versa.      However, Tibet is not entirely situated at a migration crossroad of the floral ele- ments. An ample amount of the data shows that Compositae flora have a particular capability of development in Tibet. of the total number of species of Tibetan Com- positae, 102 species and 1 genus (Ajaniopsis Shih) are endemic. Besides, 8 genera are re- gional endemics with their range extending to its neighbourhood. The higher percentage of endemics at specific level than at generic in Tibetan Compositae may be a result of active speciation in response to the new enviromental conditions created by the uplifting of the Himalayas.  The flora in Tibetan Plateau as a whole appears to be of a younger age.       3) The uprising of the Himalayas and of the Tibetan Plateau accompanied by the ultraviolet ray radiation, the microthermal climate and the high wind pressure has, no doubt, played a profound influence upon the speciation of the native elements of Tibetan Compositae. The recent speciation is the main trend in the development of the Com-positae flora native in Tibet in the wake of upheaval of the plateau.     

中国种子植物特有属的数量分析

Chinese flora with many endemic elements is highly important in the world’s flora. According to recent statistics there are about 196 genera of spermatophytes, be- ing 6.5% of total Chinese genera.  These endemic genera comprising 377 species belong to 68 families, among which the Gesneriaceae (28 genera), Umbelliferae (13), Compo- sitae (13), Orchidaceae (12) and Labiatae (10) are predominant.  The tropical type containing 24 families and 80 genera is dominant. After it follows the temperate type with 23 families and 50 genera.  There are also 4 families endemic to China, i.e. Gin- kgoaceae, Bretschneideraceae, Eucommiaceae and Davidiaceae.  It shows that genera endemic to China are obviously related to the tropical and temperate flora in essence.      The endemic monotypic genera (139) and endemic obligotypic genera (48) combin- ed make up more than 95% of the total number of genera endemic to China.  Phylo- genetically more than half of them are ancient or primitive.  The life forms of all ende- mic genera are also diverse.  Herbs, especially perennial herbs, prevail with the propor- tion of about 62%, and trees and shrubs are the next, with 33%, and the rest are lianas.       Based upon the calculated number of genera endemic to China in each province and the similarity coefficents between any two provinces, some conclusions may be drawn as follows:       Yunnan and Sichuan Provinces combined are the distribution centre of genera en- demic to China and may be their original or  differentiation area,  because  numerous endemic genera, including various groups, exist in these two provinces.  The second is Guizhou where there are 62 endemic genera.  Others form a declining order, south China, central China and east China. But towards the north China endemic genera de- crease gradually, and the Qinling Range is an important distributional limit.       The largest simitarity coefficient, over 50%, appears between Shaanxi and Gansu probably because of the Qinling Range linking these two provinces.  But between any other two provinces it is less than 30% and it is generaly larger between two south pro- vinces than between two north provinces.       These characteristics mentioned above are correlated with topography and climate, and they may be resulted from the diversification in geography and climatic influence for a long time.     

木兰科分类系统的初步研究

A new system of classification of Magnoliaceae proposed.  This paper deals mainly with taxonomy and phytogeography of the family Magnoliaceae on the basis of external morphology, wood anatomy and palynology.  Different  authors have had different ideas about the delimitation of genera of this family, their controversy being carried on through more than one hundred years (Table I).  Since I have been engaged in the work of the Flora Reipublicae Popularis Sinicae, I have accumulated a considerable amount of information and material and have investigated the living plants at their natural localities, which enable me to find out the evolutionary tendencies and primitive morphological characters of various genera of the family.  According to the evolutionary tendencies of the characters and the geographical distribution of this family I propose a new system by dividing it into two subfamilies, Magnolioideae and Liriodendroideae Law (1979), two tribes, Magnolieae and Michelieae Law, four subtribes, Manglietiinae Law, Magnoliinae, Elmerrilliinae Law and Micheliinae, and fifteen genera (Fig. 1 ), a system which is different from those by J. D. Dandy (1964-1974) and the other authors.      The recent distribution and possible survival centre of Magnoliaceae. The members of Magnoliaceae are distributed chiefly in temperate and tropical zones of the Northern Hemisphere, ——Southeast Asia and southeast North America, but a few genera and species also occur in the Malay Archipelago and Brazil of the Southern Hemisphere. Forty species of 4 genera occur in America, among which one genus (Dugendiodendron) is endemic to the continent, while about 200 species of 14 genera occur in Southeast Asia, of which 12 genera are endemic.  In China there are about 110 species of 11 genera which mostly occur in Guangxi, Guangdong and Yunnan; 58 species and more than 9 genera occur in the mountainous districts of Yunnan.   Moreover,  one  genus (Manglietiastrum Law, 1979) and 19 species are endemic to this region.  The family in discussion is much limited to or interruptedly distributed in the mountainous regions of Guangxi, Guangdong and Yunnan.  The regions are found to have a great abundance of species, and the members of the relatively primitive taxa are also much more there than in the other regions of the world.      The major genera, Manglietia, Magnolia and Michelia, possess 160 out of a total of 240 species in the whole family.  Talauma has 40 species, while the other eleven genera each contain only 2 to 7 species, even with one monotypic genus.   These three major genera are sufficient for indicating the evolutionary tendency and geographical distribution of Magnoliaceae.  It is worthwhile discussing their morphological  characters  and distributional patterns as follows:      The members of Manglietia are all evergreen trees, with flowers terminal, anthers dehiscing introrsely, filaments very short and flat, ovules 4 or more per carpel.  This is considered as the most primitive genus in subtribe Manglietiinae.  Eighteen out of a total  of 35 species of the genus are distributed in the western, southwest to southeast Yunnan. Very primitive species, such as Manglietia hookeri, M. insignis  and M. mega- phylla, M. grandis, also occur in this region. They are distributed from Yunnan eastwards to Zhejiang and Fujian through central China, south China, with only one species (Manglietia microtricha) of the genus westwards to Xizang.  There are several species distributing southwards from northeast India to the Malay Archipelago (Fig. 7).      The members of Magnolia are evergreen and deciduous trees or shrubs, with flowers terminal, anthers dehiscing introrsely or laterally, ovules 2 per carpel, stipule adnate to the petiole.  The genus Magnolia is the most primitive in the subtribe Magnoliinae and is the largest genus of the family Magnoliaceae. Its deciduous species are distributed from Yunnan north-eastwards to Korea and Japan (Kurile N. 46’) through Central China, North China and westwards to Burma, the eastern Himalayas  and northeast India.  The evergreen species are distributed from northeast  Yunnan  (China)  to  the Malay Archipelago.  In China there are 23 species, of which 15 seem to be very primi- tive, e.g. Magnolia henryi, M. delavayi, M. officinalis and M. rostrata, which occur in Guangxi, Guangdong and Yunnan.      The members of Michelia are evergreen trees or shrubs, with flowers axillary, an- thers dehiscing laterally or sublaterally, gynoecium stipitate, carpels numerous or few. Michelia is considered to be the most primitive in the subtribe Micheliinae, and is to the second largest genus of the family.  About 23 out of a total of 50 species of this genus are very primitive, e.g. Michelia sphaerantha, M. lacei, M. champaca,  and  M. flavidiflora, which occur in Guangdong, Guangxi and Yunnan (the distributional center of the family under discussion)  and extend eastwards to Taiwan  of  China, southern Japan through central China, southwards to the Malay Archipelago through Indo-China. westwards to Xizang of China, and south-westwards to India and Sri Lanka (Fig. 7).      The members of Magnoliaceae are concentrated in Guangxi, Guangdong and Yunnan and radiate from there.  The farther away from the centre, the less members we are able to find, but the more advanced they are in morphology.  In this old geographical centre there are more primitive species, more  endemics  and  more monotypic genera. Thus it is reasonable to assume that the region of Guangxi, Guangdong and Yunnan, China, is not only the centre of recent distribution, but also the chief survival centreof Magnoliaceae in the world.     

论胡桃科植物的地理分布

 The present paper aims to discuss the geog raphical distribution of the Juglandaceae on the basis of unity of the phylogeny and the process of dispersal in the plants.       The paper is divided into the following three parts:       1.  The systematic positions and the distribution patterns of nine living genera in the family Juglandaceae (namely, Engelhardia, Oreomunnea, Alfaroa, Pterocarya, Cyclo- carya, Juglans, Carya, Annamocarya and Platycarya) are briefly discussed.  The evolu- tional relationships between the different genera of the Juglandaceae are elucidated. The fossil distribution and the geological date of the plant groups are reviewed.  Through the analysis for the geographical distribution of the Juglandaceous genera, the distribu- tion patterns may be divided as follows:       A.  The tropical distribution pattern       a. The genera of tropical Asia distribution: Engelhardia, Annamocarya.       b. The genera of tropical Central America distribution: Oreomunnea, Alfaroa.       B.  The temperate distribution pattern       c. The genus of disjunct distribution between Western Asia and Eastern Asia: Pterocarya.       d. The genus of disjunct distribution between Eurasia and America: Juglans.      e. The genus of disjunct distribution between Eastern Asia and North America: Carya.      f. The genera whose distribution is confined to Eastern Asia: Cyclocarya, Platy- carya.      2.  The distribution of species      According to Takhtajan’s view point of phytochoria, the number of species in every region are counted.  It has shown clearily that the Eastern Asian Region and the Coti- nental South-east Asian Region are most abundant in number of genera and species. Of the 71 living species, 53 are regional endemic elements, namely 74.6% of the total species. The author is of the opinion that most endemic species in Eurasia are of old endemic nature and in America of new endimic nature.  There are now 7 genera and 28 species in China, whose south-western and central parts are most abundant in species, with Pro- vince Yunnan being richest in genera and species.      3.  Discussions of the distribution patterns of the Juglandaceae A.       The centre of floristic region B.         The centre of floristic regions is determined by the following two principles:  a. A large number of species concentrate in a district, namely the centre of the majority; b.  Species of a district can reflect the main stages of the systematic evolution of the Juglandaceae, namely the centre of diversity.  It has shown clearly that the southern part of Eastern Asian region and the northern part of Continental South-east Asian Region (i.c. Southern China and Northern Indo-China) are the main distribution centre of the Juglandaceae, while the southern part of Sonora Region and Caribbean Region  (i.c. South-western U.S.A., Mexico and Central America) are the secondary distribution centre.       As far as fossil records goes, it has shown that in Tertiary period the Juglanda- ceae were widely distributed in northern Eurasia and North America, growing not only in Europe and the Caucasus but also as far as in Greenland and Alaska.  It may be considered that the Juglandaceae might be originated from Laurasia.  According to the analysis of distribution pattern for living primitive genus, for example, Engelhar- dia, South-western China and Northern Indo-China may be the birthplace of the most primitive Juglandaceous plants.  It also can be seen that the primitive genera and the primitive sections of every genus in the Juglandaceae have mostly distributed in the tropics or subtropics. At the same time, according to the analysis of morphological cha- racters, such as naked buds in the primitive taxa of this family, it is considered that this character has relationship with the living conditions of their ancestors.  All the evidence seems to show that the Juglandaceae are of forest origin in the tropical moun- tains having seasonal drying period.       B.  The time of the origin       The geological times of fossil records are analyzed. It is concluded that the origin of the Juglandaceae dates back at least as early as the Cretaceous period.       C.  The routes of despersal       After the emergence of the Juglandaceous plant on earth, it had first developed and dispersed in Southern China and Indo-China.  Under conditions of the stable tempera- ture and humidity in North Hemisphere during the period of its origin and development, the Juglandaceous plants had rapidly developed and distributed in Eurasia and dis- persed to North America by two routes: Europe-Greenland-North  America  route  and Asia-Bering Land-bridge-North America route.  From Central America it later reached South America.      D.  The formaation of the modern distribution pattern and reasons for this forma- tion.      According to the fossil records, the formation of two disjunct areas was not due to the origin of synchronous development, nor to the parallel evolution in the two con- tinents of Eurasia and America, nor can it be interpreted as due to result of transmis- sive function.  The modern distribution pattern has developed as a result of the tectonic movement and of the climatic change after the Tertiary period.  Because of the con- tinental drift, the Eurasian Continent was separated from the North American Conti- nent, it had formed a disjunction between Eurasia and North America. Especially, under the glaciation during the Late Tertiary and Quaternary Periods, the continents in Eu- rasia and North America were covered by ice sheet with the exception of “plant refuges”, most plants in the area were destroyed, but the southern part of Eastern Asia remained practically intact and most of the plants including the Juglandaceae were preserved from destruction by ice and thence became a main centre of survival in the North Hemisphere, likewise, there is another centre of survival in the same latitude in North America and Central America.      E.  Finally, the probable evolutionary relationships of the genera of the Juglanda-ceae is presented by the dendrogram in the text.     

东喜马拉雅南翼苔藓植物区系的特性及其来源

 In the south-east and south Xizang, in cluding Medog, Zayü some western separate valleys Yadong, Kama near Zentang in Dinggye, Boqu near Zham in Nyalam and Gyirong, a mild climate prevails because of the very high mountains and the very deep valleys.  According to our preliminary survey, 4/5 of the genera and 7/10 of the species, i.e. approximately representing all families and genera of the tropical and subtropical bryofliora of Xizang, are restricted to these localities below the altitude of 2,300 meters.  It almost agrees with the previous presumption that the Tsangpo gorge is the line of connection between two paleoeontinents—Laurasia and Gondwana.       Moreover, the bryoflora of these localities, besides the Indo-Malasian elements and East Asian elements as the main components, has at least about 40 genera in common with south America, Australia and Africa.  According to the historical phytogeogra- phical point of view, the distribution range of centain genera is formed through a period of long historical development.  The same is true for the area of different species, although they are found in widely separate areas right now, yet they might have once a continuous distribution in certain historical age. The Indian plate collided against the eastern part of Laurasia and afterwards the Australasian plate moved to the north.  All these might have dispersed the Gondwana elements as far as to the southeastern part of Xizang.      It is very interesting to note that of the 32 genera of bryophytes endemic to East Asia, 13 have recently been found in the southeast and south Xizang and also in the neighbouring regions, i.e. Yunnan, Sichuan, where there are many genera being in common with southeast and south Xizang and also highly concentrated in distribution. This may suggest that the Himalayas, being the highest and youngest mountain range, have changed the atmospheric circulation, and have created a new ecological condition between tropical and frigid zones, which have given the distribution of the newly form- ed genera a suitable circumstance to survive.  It may be presumed that the region covering counties Medog, Zayü, Yadong etc. in southeastern and southern parts of the Himalayas is a new center of distribution of bryophytes under the influence of the up-heaval of the Himalayas.     

白洋淀水生植物区系初步分析

The Baiyandian Lake lies at 115°56′ E. and 38°55′ N.  It is about 8 m above sea level, with an area about 312 square km.  There are abundant aquatic plants here, inc- luding flowering plants of 30 families, 46 genera, 66 species and 3 varieties besides one in- troduced species.  32 species are erect ones covering about 48.5% of the total number of species in this lake, 20 are submerged ones, about 30.3%, 9 are leaf-floating ones, about 13.6% and 5 are floating ones, 7.6%.  The soil at the bed of the lake is very fertile and the depth is appro- priate, favouring various plants.  The coverage is large, sometimes it may reach 100%, for example, in communities of Phragmites communis, Hydrilla verticillata,  Hydrocharis dubis, Trapa bicornis, etc. Although the Baiyandian Lake is located in the temperate zone, the floristic elements are complex.  There are tropic-subtropic elements and elements of high-frigid zone as well.  When compared respectively with the floras of the Lakes Jingbo Lake, Honghu Lake and Lugu Lake, the flora of this lake is found more similar to that of subtropic Honghu Lake than to those of the other lakes, which shows that it hase a character transitional between the south and the north in China.  Further studies of the floristic relationships are needed in order to exploit and use the resources of the aquatic plants in this lake.     

微孔草属的研究

 1.  Having analyzed the external morphology of the genus Microula, the author has proposed a series of criteria as bases for the construction of a classification scheme of this genus.  The most important ones are as follows:      1)  The normally developed stem is primitive, and the strongly abbreviated stem more advanced.      2)  The small inconspicuous bracts are more primitive than the large suborbicular densely arranged ones, which almost entirely cover the flowers and the fruits.      3)  Nutlets with small dorsal pit  are more primitive than those with larger pit on one hand or those without it on the other.      4)  The dorsal pit with simple margin precedes that with double margins.      5)  Nutlets with subbasal areola precede those with lateral or apical areola.      6)  Nutlets without glochids precede those with glochids.      2.  Basing upon these criteria the genus Microula may be divided into six sections. The section Schistocaryum may be the primitive one, and the others may be evolved from it respectively.  The possible affinities between them are demonstrated in figure no. two.      3.  The genus Microula, containing 30 species,  is  mainly  distributed  in  the Chinghai-Tibetan plateau and the majority of its species concentrates in the eastern border of the plateau, and of the 30 species 26—that is 90 percent—are endemic to China, and the remaining 4 are distributed elsewhere in China, too, and extending southward and westward to Bhutan, Sikkim, Nepal and Kashmir respectively.  In the region between Heishui, Province Szechuan, and Chinghai Lake there are 9 species, which, curiously, represent all the six sections of Microula, hence this region seems to be the center of maximum variation of this genus.  M. ovalifolia whose nutlets have small dorsal pit and subbasal areola may be considered the most primitive species. Thus the author is of the opinion that the western part of province Szechuan, to which M. ovalifolia is endemic, may probably be the center of origin of the genusMicroula.     

燕辽地区侏罗纪杉科一新属——燕辽杉

  A fossil plant, Yanliaoia sinensis gen. et sp. nov. of late Jurassic is described from western Liaoning.  It is an arborescent conifer, deciduous, with branchlets and leaves resemble closely those of Sequoia sempervirens.. The form and structure of female cones and cone scales are similar to those of extinct genus Elatides and living genera Cunninghamia, Taiwania and Athrotaxis,  The female cone, however, is smaller.  Cone scales are spirally arranged, woody, thin and ellipticzl, having a short spine at their apex.  Seeds, globuloidal, without wings, about ten on each scale, and longitudinally arranged in three to four ranks, which occupy nearly the whole area of the ventral surface of ovuliferous scales.      The author believes that Yanliaoa and Elatides are two closely allied genera, evolved roughly in parallel with each other in the late Mesozoic.  Yanliaoa may be one of the ancestors of the living genera of Taxodiaceae. remarkablly the lineal or sublineal ascendant of Sequoia sempervirens.   The  so-called  “Sequoid jeholensis”(Endos. 1951) may be a branchlet of Yanliaoa sinensis.     

中国清风藤科的初步研究

 This paper is a preliminary study on the Sabiaceae in aspects of its morphology, taxonomy and geography.  We propose that the Sabioideae and Meliosmoideae as two new subfamilies of Sabiaceae according to the external morphology, flower structure and geographical distribution of these two genera respectively.       This paper follows the taxonomic concepts of Luetha Chen on Sabia and C. F. van Beusekom on Meliosma.  We agree with them for their classification of these two genera above the specific rank.  As to the revision work of Sabia by van de Water and C. F. van Beusekom’s work on Meliosma we disagree  for their unduly broad specific concepts.  We rather treat the species of these two genera according to their habitats in regions on a relatively narrower sense.  The genus Sabia of China are classified into 2 tribes, with 16 species, 5 subspecies and 2 varieties in which 4 sub- species and l variety are as new combinations, the genus of Meliosma in China are classified into 2 subgenera with 29 species, and 7 varieties of which 4 varieties are new combinations.       After examining the affinity of the species of Sabia and Meliosma in China and its neighboring nations such as Burma, Japan and Bhutan, we found that their migra- tion initiated from China, as the primitive species of these two genera occured in northeast and central part of Yunnan, sou theast of Sichuan, north of Guizhou and west of Hubei, the region may probably be the main origin of these two genera.      As shown in tables 1 & 2, the localities where the species of these two genera den- sely populate they are from Yunnan, Guangxi, and Guangdong coinciding with the concepts of C. F. van Beusekom and van de Water about the distribution of exotic species of these two genera, it may reasonable be pointed out that the center of distri- bution of these two genera is Yunnan, Guangxi, Guangdong and nieghboring nations, upper Burma and northern Vietnam.  Futhermore, it may be seen that starting from this center the number of species become less and less as they proceed far and far awaybut become more advance in evolution.     

三峡大老岭植物区系的垂直梯度分析

为探讨山地植物区系构成特征及其垂直梯度的生态意义，根据对三峡大老岭地区植被垂直样带 调查获得的植物区系资料，分析了该地区植物区系成分构成的基本特征及其随海拔梯度的变化趋势，寻找了区系平衡点的位置；并利用聚类方法分析了山地气候垂直分异对区系成分构成的影响。结果表明：①大老岭植物区系具有温带性质，但仍反映了与热带区系的历史联系，有强烈的区域性；②属的分布区类型可归为热带分布、温带分布、地中海—中亚中心和东亚中心4组，各组区系成分的垂直梯度特征不同；热带、亚热带成分与温带成分的平衡点大致位于海拔650m；③区系成分构成和属的物种数量构成的聚类分析结果一致显示了植物区系构成与山地气候和植被垂直带相对应的格局。     

秦岭植物区系的性质、特点和起源

 秦岭是中国长江和黄河两大水系的分水岭，位于北纬32°5′至34°45′；东经104°30′至115°52′，最高峰达3767m。该山区是我国温带植物区系最丰实的地区之一，约有种子植物3124种，隶属于158科，892属。包括热带属220属，温带属563属，和中国特有属39属。根据该山区植物区系中各大科、主要植物群落优势种和组成种类的温带性质以及温带属在整个植物区系中的主导地位。该山区的植物区系和植被具有明显的温带性特点。特有种和非特有种的分析结果表明，该山区植物区系的特点还表现在高度特有性和以中国-日本森林植物区系为主体方面。    根据古植物学资料分析，秦岭地区植物区系的起源时间不会晚于晚白垩纪；植物群落的主要成份可能以原地生长的种类为主；秦岭及其邻近古老山区，不仅对自身的植物区系和植被具有较大的发生意义，而且对东亚植物区系具有始生性质。     

中国亚热带地区植物区系地理成分及其空间格局的数量分析

选择我国亚热带区域76个地区(不包括行政单元)的植物区系调查和研究资料,采用主成分分析 (PCA)、TWINSPAN聚类和样带梯度分析方法,研究了我国亚热带区域植物区系地理成分的构成特征和 各成分之间的关系;各研究区域间植物区系地理成分构成的相关性和空间分异特征;并分析了贯穿我国亚热带区域的东经108.5°～111.5°和北纬29°～31°两条样带的几种区系地理成分的分布梯度。结果表明：①15种区系地理成分可以归为热带成分、温带成分、干旱区成分和东亚中心成分4组;我国亚热带各地区区系地理成分中泛热带和北温带分布的比例最高;干旱区分布诸类比例极小;东亚中心成分的比例突出,特有性强;②样点的PCA排序明显反映了北、中、南亚热带在区系地理成分构成上的差异,但亚热带东西部的差异不明显;③样带分析清晰地反映了热带、温带成分之比的梯度格局;川东鄂西地区特有分布中心的地位和世界分布与特有成分相反的分布格局。     

峨眉山兰科植物的地理分布和区系特点

 Situated in western part of Sichuan Province, 29°30'N, 103°20'E, the sacred Mt. Emei is one of the well-known large mountains in China. Its summit is about 3100 m ab- ove sea level with a relative height of 2550 m.        The orchid flora in Mt. Emei so far known comprises 47 genera and 109 species, among which 21 are epiphytes, 83 terrestrials and 5 saprophytes (Table 1.)        1.  The vertical distribution of the orchid flora in the mountain. The epiphytic orchids are concentrated in the lower region  below Hongchunping and Wanniansi (1100 m alt.), where there are 20 species, which make over 95% of epiphytic species; the upper limit for the epiphytic orchids is Jiulaodong and Chudian ( 1800 m alt.). The terrestrial orchids also mainly occur at the lower region below Jiulaodong and Chudian (1800 m alt.), where there are 54 species, most of which are found at even lower part of the mountain, below Hongchunping and Wanniansi (1100 m alt.). The tropicas orchids in the mountain, such as Cleisostoma, Vanda, Holcoglossum, Tropiclia, Thunia, Mischobulbum, Ludisia, Anoectochilus, Odontochilus, etc. all grow only at the lower part of the mountain below Hongchunping and Wanniansi (1100 m alt.).        2. The floristic features of the orchid flora in the Mt. Emei.        (1)  The orchid flora in the mountain so far known comprises 47 genera (over 2/3 of the total orchid genera in Sichuan) and 109 species (over 1/3 of the total orchid species in Sichuan). The Mt. Emei is very rich in orchid species, as compared with neighbouring mountains of same magnitude, such as Mt. Shennonjia in western Hubei, Qin Ling in sou- thern Shaanxi, Jinfo Shan in south-eastern Sichuan, and Erlang Shan in western Sichuan.        (2)  The orchids in the mountain are complex in floristic components as indicated below:        1)  Twenty seven species, belonging to 18 genera, are widespread, covering the whole East-Asian region.        2)  Twenty three species, belonging to 15 genera, are the elements of the Sino-Japanese Subregion. Among them 13 species occur only in Japan and eastern China with the mountain ar the westernmost limit, but the other species extend westwards as far as Kangding and ErLang Shan or Baoxing in Sichuan Province.        3)  Forty two species, belonging to 22 genera, are the elements of the Sino-Himala-yan Subregion, with 5 species having their range extending from the Himalayan  region eastwards to Mt. Emei.        4)  Some tropical genera (8 species), belonging to Indo-Malaysian floristic elements, have the mountain as their northern limit of distribution.       The orchid flora of the Mt. Emei contains not only the East-Asian elements, but also some Indo-Malaysian elements, though its composition is mainly of the temperate and subtropical Eastern Asian (Sino-Japanese) ones.        (3)  The orchid flora in the mountain is characterized by geographical vicariation and differentiation.       There are nine species-pairs (belong to genera Calanthe,  Platanthera,  Dendrobium etc.) of the vertical vicarism and six species-pairs (belonging to genera Tropidia, Aneoctochilus, Mischobulbum, Gymnadenia Orchis, etc.) of the horizontal vicarism in the Mt.  Emei.       Remarkable differentiation of orchid flora in the Mt. Emei is shown in the abundance of endemic elements and as clear geographical vicariation.        (4)  There are 8 endemic species and one variety of orchids in the Mt. Emei, more  abundant than in Xizang.      The floristic features of the orchid flora of the Mt. Emei are rich in species, compara tively complex in components, rather prolific in endemic species, and characterized by geographical vicariation and differentiation. The orchid flora in the Mt. Emei mainly consists of the subtropical and temperate East-Asian elements, with a  considerable proportion of tropical elements though.     

秦岭太白山地区的植物区系和植被

太白山位居秦岭的中段，为秦岭山脉第一高峰。该山区是我国温带植物区系最丰富的地区之一， 约有种子植物1782种，隶属于125科，657属，包括热带属130属，温带属436属和特有属24属。该地 区20个较大科的种数，约占其全部植物区系的66.6％，其中特有种653种。本文对这些大科的性质及 其在植物区系和植被中的作用进行了分析讨论。在分析了全部属和植物群落优势种的地理分布的基础 上，着重讨论了该地区与其他九个山区的植物区系关系。   　本文对植被垂直带划分；主要植物群落的基本特点和区系相似性；物种多样性与海拔高度变化的关 系以及生活型谱与不同坡向和海拔变化的关系等作了具体分析。对该地区主要植物群落的形成时期以 及对秦岭地区植物区系的过渡性质提出了自己的看法。从植物学角度出发，秦岭地区作为亚热带和温带 植物的分界线的提法与该地区的植物区系和植物群落性质以及水热条件情况不相符合     

中国及其邻近地区松杉类特有属的现代生态地理分布及其意义

 1．我国及其邻近地区松杉类特有属，主要分布于我国东南部、南部和西南 部，大约相当于我国亚热带常绿阔叶林带的范围。其垂直分布一般在海拔100— 1800米之间，少数属可达2800米，但不逾越海拔3000米。 2．我国松杉类特有属分布地区的水热条件，大致为年平均温度在10℃-20℃之间，绝对最低温度为-6.3℃——11.3℃， 年降水量一般在2000毫米左右。土壤pH 4．0—5．5之间，呈酸性反应。 3．我国及其邻近地区松杉类特有属数约占全世界松杉类特有属数的37.5%，是世界上最丰富、分布最为集中的地区。这些属的化石出现于晚白垩纪或第三纪时期。  因此，我国无疑是松杉类特有属的现代地理分布中心和保存中心。这对进一步研究松杉类植物的发生和发展，具有重要的意义。     

重楼属植物的细胞地理学研究

重楼属Paris有19种，分布于欧亚大陆。根据对本属所有种的染色体研究，重楼属染色体基数为5，核型的基本结构有两种形式：热带核型K2n＝2x=10=6m十4t和温带核型K2n=2x=10＝6m+4st或  6m+2st十2t。  热带核型的种(13种)分布在亚洲大陆的热带和亚热带；温带核型的种(6种)则出现在欧亚大陆的温带地域。重楼属的多倍体种的核型属温带核型，为本属的边陲种。四倍体种四叶重楼Paris quadrifolia分布在本属分布区的西端(欧洲)；日本重楼P.japonica是八倍体种，局限在属分布区的东端(日本)。全部热带核型的种都是二倍体种。其中海南重楼P.dunniana等较原始种类都集中在华南和中南半岛北部。作者认为，亚洲大陆北纬18°至北回归线的热带地域是重楼属的起源地，云贵高原至邛崃山地域拥有14种重楼和9种核型结构式(全属有13种核型结构式)，是重楼属的多样化中心，即现代分布中心。     

试论中国种子植物特有属的分布区类型

中国种子植物特有属是局限分布于中国行政区域范围内的植物成分，就其分布特点看，集中分布于中国南部亚热带广阔区域。由于中国地域广袤，虽然大多数特有属分布在东亚自然地域范围内，但南部特有属的分布范围已进入古热带植物区的马来亚森林植物亚区的北部，而西部的特有属的分布范围已进入青藏高原地区。局限于不同地域分布的特有属，各自的起源发生、所经历的地质历史过程存在一定差别。本文以自然地理区划作为研究中国种子植物特有属分布区类型的依据，将中国特有属分布区类型划分为中国东部和中部特有分布变型、中国南部特有分布变型、中国西部特有分布变型和中国北部特有分布变型4类。其中中国南部特有分布变型所含特有属为热带区系成分，其它3个特有分布变型所含特有属为温带区系成分。这样能较客观地反映中国特有属的自然地理特征，有利于研究局部地区植物区系的地质历史演变过程。